Nola internella-analis complex NA -

Taxonomic Information

Nola analis (Wileman & West, 1928), Nola internella (Walker, 1864), Nola pascua (Swinhoe, 1885), Nola quadrimaculata Heylaerts, 1892. These individuals could be any of these four species, all of which have a similar fascia and are found in India. They are best separated by examining specimens and/or genitalia.

List of Nola Leach, 1815 known from India (vide Joshi et al. 2019):

Nola analis (Wileman and West, 1928) (Celema). India: South India; Sri Lanka; Nepal.
Nola angulata (Moore, 1888) (Roeselia). India: NW Himalayas, Himachal Pradesh, Tamil Nadu; Sri Lanka.
Nola astigma Hampson, 1894. India: (West Bengal, Sikkim, Manipur, Nagaland.
Nola atrocincta Inoue, 1998. India: Sikkim; Nepal.
Nola cingalesa (Moore, 1882). India: Tamil Nadu, Nilgiris; Sri Lanka.
Nola cretacea (Hampson, 1901) (Celama). India: Maharashtra, West Bengal, South India; Nepal.
Nola defreina László, Ronkay and Witt, 2010. India: Assam, Meghalaya, Eastern Himalaya; Central Nepal.
Nola duplicilinea (Hampson, 1900) (Celama). India: Sikkim, NE Himalaya; Nepal.
Nola endotherma (Hampson, 1918) (Celama). India: Meghalaya.
Nola erythrostigmata Hampson, 1894. India: NE Himalaya, Assam; Bhutan.
Nola euryzonata (Hampson, 1900) (Celama). India: Mizoram.
Nola fasciata (Walker, 1866) (Minnagara). India: Sikkim, Nilgiris, Tamil Nadu, NE Himalaya; Sri Lanka; Nepal.
Nola fraterna (Moore, 1888) (Roeselia). India: Himachal Pradesh, NW Himalaya, Himalaya, Andaman Is.
Nola fuscantea (van Eecke, 1920) (Celama). India: Sikkim, West Bengal; Nepal.
Nola infralba Inoue, 1976. India: Arunachal Pradesh.
Nola internella (Walker, 1864) (Pisara). India: West Bengal, Maharashtra, Nilgiris; Bhutan; Sri Lanka.
Nola lucidalis (Walker, 1864 [1865]) (Pisara). India: Sikkim, Indian Subregion; Sri Lanka.
Nola marginata Hampson, 1895. India: Sikkim, Arunachal Pradesh, Uttarakhand, Himalaya; Bhutan.
Nola mesomelana (Hampson, 1900) (Celama). India: Sikkim, NE Himalaya.
Nola mesosticta (Hampson, 1900) (Neonola). India: Sikkim, Indian Subregion; Sri Lanka.
Nola negrita Hampson, 1894. India: Karnataka, Tamil Nadu; Sri Lanka.
Nola ochrolopha (Hampson, 1911) (Celama). India: Sikkim, NE Himalaya.
Nola pascua (Swinhoe, 1885) (Roeselia). India: Tamil Nadu, Nagaland, Maharashtra; Sri Lanka.
Nola polia Hampson, 1900 (Celama). India: Sikkim.
Nola pumila Snellen, 1875. India: Sikkim, Assam, Meghalaya, Maharashtra, Nilgiris.
Nola quadrimaculata Heylaerts, 1892. India: NE Himalaya, South India.
Nola sikkima (Moore, 1888) (Roeselia). India: Sikkim, West Bengal; East Nepal.
Nola spinivesica Holloway, 2003. India.
Nola spreta Butler, 1880. India: Nilgiris, Nagaland, Assam. 
Nola suffusa (Hampson, 1900) (Celama). India: Sikkim.
Nola taeniata Snellen, 1875. India: Punjab, West Bengal, Indian sub-region; Sri Lanka; Nepal; Myanmar.
Nola thyrophora (Hampson, 1914) (Pisara). North India; Bhutan; Myanmar.
Nola wernerthomasi László, Ronkay and Witt, 2010. India: Sikkim, Assam, Meghalaya; Nepal.

Photo Gallery and Species Biology

StateJanFebMarAprMayJunJulAugSepOctNovDecNo date
Andaman and Nicobar Islands
Andhra Pradesh1
Arunachal Pradesh
Assam1
Bihar
Chandigarh
Chhattisgarh1
Dadra & Nagar Haveli
Daman & Diu
Delhi11
Goa1
Gujarat5421212
Haryana
Himachal Pradesh
Jammu and Kashmir UT
Jharkhand
Karnataka11412
Kerala1
Ladakh UT
Lakshadweep
Madhya Pradesh
Maharashtra2411
Manipur
Meghalaya
Mizoram
Nagaland
Odisha
Paschimbanga
Pondicherry
Punjab
Rajasthan1
Sikkim
Tamil Nadu
Telangana
Tripura
Uttar Pradesh1
Uttarakhand1111
West Bengal11
Total6421112361085
Nola internella-analis complex: Cajanus cajan (Fabaceae), Durio zibethinus (Malvaceae), Cenchrus americanus (Poaceae), Rubus spp. (Rosaceae), Terminalia paniculata (Combretaceae) (Robinson et al. 2010). 
Nola analis (Wileman & West, 1928), Nola internella (Walker, 1864), Nola pascua (Swinhoe, 1885), Nola quadrimaculata Heylaerts, 1892.
All these four species have a double triangular spot on the forewing costa and a double postmedial line on the forewing. They are best separated by examining specimens and/or genitalia. Named as the internella-analis complex as Nola internella (Walker, 1864) was the first of these species to be described and Nola analis (Wileman & West, 1928) is possibly the most widespread species.

See references below: 
  1. https://www.mothsofborneo.com/part-18/nolinae/nolinae_14_8.php 
  2. https://www.inaturalist.org/taxa/244416-Nola-analis for analis
  3. https://www.inaturalist.org/taxa/199523-Nola-quadrimaculata for quadrimaculata
  4. https://www.inaturalist.org/taxa/1115039-Nola-pascua for pascua
  5. Joshi, R., Singh, N. and Laszlo, G.M., 2019. An updated Checklist of subfamily Nolinae (Lepidoptera, Nolidae) from India. Records of the Zoological Survey of India, 119(3), pp.263-281. Lists Nola analis, quadrimaculata, pascua and internella from India. 
 

Page citation

Anonymous 2026. Nola internella-analis complex NA – . In Sondhi, S., R.P. Singh, G. Iyer, J. D'silva and K. Kunte (Chief Editors). Butterflies of India, v. 4.11. Published by the Indian Foundation for Butterflies. URL: https://www.mothsofindia.org/nola-internella-analis-complex, accessed 2026/02/15.