Glyphodes pyloalis complex

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sanjay.sondhi1@gmail.com
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Suggested ID: Glyphodes pyloalis complex Walker, 1859
Glyphodes pyloalis Walker, 1859. List Spec. Lepid. Insects Colln Br. Mus. 19: 973. TL: N. China. Text at https://archive.org/details/listofspecimenso1921brit/page/973/mode/1up?view=theater
Glyphodes sylpharis Butler, 1878. Ill. typical Spec. Lepid. Het. Colln Br. Mus. 2: 57, pl. 39, f. 2. TL: Yokohoma. Text at https://archive.org/details/illustrationsoft02brit/page/57/mode/1up?view=theater. Plate at https://archive.org/details/illustrationsoft02brit/page/n114/mode/1up?view=theater
http://mothphotographersgroup.msstate.edu/species.php?hodges=5197
https://www.butterfliesandmoths.org/species/Glyphodes-pyloalis
https://v3.boldsystems.org/index.php/Taxbrowser_Taxonpage?taxid=273239
http://bugguide.net/node/view/20378
Listed in Mathew’s Pyralid paper.
Hampson vol 4, pg 357.
Roger Kendrick PhD Plate 14/33
Sondhi et al 2021. Moths of Tale. Pl. 2/34.
https://www.inaturalist.org/taxa/154624-Glyphodes-pyloalis

There are two other species in this complex that are similar:
Glyphodes harutai Yamanaka, 1998: TL: Nepal
Glyphodes duplicalis Inoue, Munroe & Mutuura, 1981. TL: Japan.

Glyphodes harutai Yamanaka, 1998. Pyralidae of Nepal (II). In: Moths of Nepal Part 5, TINEA. Pg. 108, pl. 142/15.
Expanse 21-23mm. Very similar to G. duplicalis Inoue, Munroe & Mutuura, 1981, Tinea 11: 91, figs 1-2, 5, 7, described from Japan in colour and maculation, but ground colour of upperside of both wings more or less strongly tinged with buff; orbicular spot a little smaller; subterminal and terminal band of hindwing a little narrower.
Male & female genitalia (Figs 892, 895). Very similar to G. pyloalis Walker, 1859 described from north China rather than to G. duplicalis, but different from those of the former as follows. In male apico-dorsal portion of the uncus more weakly dilated (Fig. 893). In female collar of ductus bursae a little shorter and more or less strongly sclerotized.
Moth of Nepal, Part 5, pg.
Irungbam et al, Fig 5/E for Glyphodes harutai.
https://v3.boldsystems.org/index.php/Taxbrowser_Taxonpage?taxid=1143516 for harutai.
Most Indian literature including Singh et al 2022. A catalogue of Indian Pyraloidea (Lepidoptera), treats this as Glyphodes pyloalis. But the TL of pyloalis is N. China. So the entire Indian population might be Glyphodes harutai (TL: Nepal), if this is a valid taxon! I think this all needs resolution, but for now, lumping this into Glyphodes pyloalis complex seems the way to go.

GI: Glyphodes harutai. See remarks in 231097, 237458
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Sanjay Sondhi
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MOI-pnz200
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Butterfly
Moth Taxon search:
Month
August
Year
2019
Day
31